Pasting returns/newlines as spaces in 'Save as' dialog
A few versions back (somewhere around 35–36) pasting text with carriage returns/newlines into the File name box would cause those returns/newlines to turn into space characters. I prefer this over the current behavior where only the text before the first return/newline is pasted.
It seems that the "editor.singleLine.pasteNewlines" preference (which I have set to "2") isn't being obeyed.
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Where to go for developer support
I would also highly recommend filing a bug with an example in bugzilla.mozilla.org.
editor.singleLine.pasteNewlines only applies to single line input fields on a web page, so that won't help in your case.
I assume that spaces should work although you might end up with %20 in some cases, but I don't know about other whitespace characters. It would normally be best to use underscore characters instead of whitespace characters in file names.